Phengodidae

Phengodidae
	Phengodes orbignyi
	Taxonomische indeling
Rijk:	Animalia (Dieren)
Stam:	Arthropoda (Geleedpotigen)
Klasse:	Insecta (Insecten)
Orde:	Coleoptera (Kevers)
Onderorde:	Polyphaga
Infraorde:	Elateriformia
Superfamilie:	Elateroidea
	Familie
	Phengodidae; LeConte, 1861
	Afbeeldingen op Wikimedia Commons
	Phengodidae op Wikispecies
Portaal	Biologie ; Insecten

Phengodidae zijn een familie van kevers. De familie is voor het eerst wetenschappelijk beschreven in 1861 door LeConte. Het is een van de families van glimwormen.

Taxonomie[bewerken | brontekst bewerken]

De volgende taxa zijn bij de familie ingedeeld:^[1]

Onderfamilie Phengodinae LeConte, 1861
- = Pseudophengodidae Pic, 1930
- Geslacht Microphengodes Wittmer, 1976
  - Microphengodes howdeni Wittmer, 1976
  - Microphengodes longicornis Wittmer, 1976
- Geslacht Phengodes Illiger, 1807
  - = Astraptor Murray, 1868
  - Ondergeslacht Phengodella Wittmer
    - Phengodes atezcanus Zaragoza, 1981
    - Phengodes bella Barber, 1913
    - Phengodes bimaculata Gorham, 1881
      - = Phengodes nevermani Pic, 1937
      - = Phengodes notaticollis Pic, 1937:
    - Phengodes bipennifera Gorham, 1881
      - = Phengodes brasiliensis Pic, 1925:
    - Phengodes brailovskyi Zaragoza & Wittmer, 1986
    - Phengodes championi Pic, 1927
    - Phengodes ecuadoriana Witmer, 1988
    - Phengodes fenestrata Wittmer, 1975
    - Phengodes frontalis LeConte, 1881
    - Phengodes insignis Bourgeois, 1888
    - Phengodes insulcata Pic, 1925
      - = Phengodes atricolor Pic, 1937
    - Phengodes leonilae Zaragoza & Wittmer, 1986
    - Phengodes minor Gorham, 1881
    - Phengodes nigricornis Gorham, 1881
      - = Phengodes bipennifera Gorham, 1881
      - = Phengodes gorhami Pic, 1927
    - Phengodes tuxtlaensis Zaragoza, 1989
    - Phengodes varicolor Zaragoza & Wittmer, 1986
    - Phengodes variicornis Zaragoza & Wittmer, 1986
    - Phengodes venezolana Wittmer, 1988
  - Ondergeslacht Phengodes Illiger, 1807
    - Phengodes arizonensis Wittmer, 1975
    - Phengodes bolivari Zarazoga, 1980
    - Phengodes fusciceps LeConte, 1861
    - Phengodes inflata Wittmer, 1975
    - Phengodes laticollis LeConte, 1881
    - Phengodes mexicana Wittmer, 1975
    - Phengodes nigromaculata Wittmer, 1975
    - Phengodes plumosa (Oliver, 1790)
      - = Lampyris plumosa Oliver, 1790
    - Phengodes vazquezae Zarazoga, 1978
- Geslacht Pseudophengodes Pic, 1930
  - Pseudophengodes bordoni Wittmer, 1988
  - Pseudophengodes brasiliensis Wittmer, 1976
  - Pseudophengodes cincinnata (Erichson, 1847)
    - = Phengodes cincinnata Erichson, 1847
  - Pseudophengodes equatoriana Wittmer, 1976
  - seudophengodes fernandezi Wittmer, 1976
  - Pseudophengodes flavicollis (Leach, 1824)
    - = Phengodes flavicollis Leach, 1824
  - Pseudophengodes floccosa (Erichson, 1847)
    - = Phengodes floccosa Erichson, 1847
  - Pseudophengodes foveiocollis Wittmer, 1976
  - Pseudophengodes fusca (Gorham, 1881)
    - = Phengodes fusca Gorham, 1881
  - Pseudophengodes gracilicornis Wittmer, 1976
  - Pseudophengodes marginata (Pic, 1925)
    - = Phengodes marginata Pic, 1925
  - Pseudophengodes marmoratus Wittmer, 1996
  - Pseudophengodes meridiana Wittmer, 1976
  - Pseudophengodes notaticollis (Pic, 1925)
    - = Phengodes notaticollis Pic, 1925
  - Pseudophengodes onorei Wittmer, 1996
  - Pseudophengodes orbignyi (Blanchard, 1837)
    - = Phengodes orbignyi Blanchard, 1837
  - Pseudophengodes penai Wittmer, 1976
  - Pseudophengodes polita Wittmer, 1976
  - Pseudophengodes pulchella (Guerin, 1843)
    - = Phengodes pulchella Guerin, 1843
  - Pseudophengodes puncticollis Wittmer, 1976
  - Pseudophengodes roulini (Guerin, 1843)
    - = Phengodes roulini Guerin, 1843
  - Pseudophengodes ruficollis (Motschulsky, 1854)
    - = Phengodes ruficollis Motschulsky, 1854
- Geslacht Zarhipis J.LeConte, 1880
  - = Zarhippus Woodworth, 1913
  - = Zarrhipis Fall, 1923
  - Zarhipis integripennis (LeConte, 1874)
    - = Phengodes integripennis LeConte, 1874
    - = Zarrhipis alamedae Fall, 1923
    - = Zarrhipis amictus Fall, 1923
    - = Zarrhipis brevicollis Fall, 1923
    - = Zarhipis piciventris LeConte, 1881
    - = Zarhipis riversi Horn, 1885
    - = Zarhipis ruficollis LeConte, 1881
  - Zarhipis tiemanni Linsdale, 1964
  - Zarhipis truncaticeps (Fall, 1923
    - = Zarrhipis truncaticeps Fall, 1923
Onderfamilie Mastinocerinae LeConte, 1881
- Geslacht Akamboja Roza, Quintino, Mermudes & Silveira, 2017^[2]
  - Akamboja caparaoensis Roza, Quintino, Mermudes & Silveira, 2017^[3]
  - Akamboja cleidae Roza, Quintino, Mermudes & Silveira, 2017^[4]
  - Akamboja minimum Roza, Quintino, Mermudes & Silveira, 2017^[5]
  - Akamboja monteirorum Roza, Quintino, Mermudes & Silveira, 2017^[6]
  - Akamboja tenebrae Roza, Quintino, Mermudes & Silveira, 2017^[7]
- Geslacht Baeoscelis Spinola, 1854
  - Baeoscelis osculata Spinola, 1854
- Geslacht Brasilocerus Wittmer, 1963
  - = Brasibcerus
  - Brasilocerus belemensis Wittmer, 1988
  - Brasilocerus dietrichi (Wittmer, 1963)
    - = Mastinocerus dietrichi Wittmer, 1963
  - Brasilocerus espiritensis Wittmer, 1963
  - Brasilocerus minasensis Wittmer, 1976
  - Brasilocerus nigerrimus Wittmer, 1996
  - Brasilocerus nigrofasciatus (Pic, 1915)
    - = Phrixothrix nigrofasciatus Pic, 1915
  - Brasilocerus oberthuri (Pic, 1955)
    - = Phrixothrix oberthuri Pic, 1955
    - = Brasilocerus impressicollis Wittmer, 1970
  - Brasilocerus opacus (Pic, 1937)
    - = Phrixothrix opacus Pic, 1937
    - = Mastinocerus callanganus Wittmer, 1963
    - = Phrixothrix subopacus Pic, 1954
  - Brasilocerus wygodzinsyi Wittmer, 1976
- Geslacht Cenophengus J.LeConte, 1881
  - Cenophengus breviplumatus Wittmer, 1976
  - Cenophengus brunneus Wittmer, 1976
  - Cenophengus ciceroi Wittmer, 1981
  - Cenophengus debilis .LeConte, 1881
  - Cenophengus gorhami Zarazoga, 1986
  - Cenophengus guerrerensis Zaragoza, 1991
  - Cenophengus howdeni Zarazoga, 1986
  - Cenophengus longicollis Wittmer, 1976
  - Cenophengus magnus Zaragoza, 1988
  - Cenophengus major Wittmer, 1976
  - Cenophengus marmoratus
  - Cenophengus pallidus Schaeffer, 1904
  - Cenophengus pedregalensis Zaragoza, 1975
  - Cenophengus punctatissimus Wittmer, 1976
  - Cenophengus villae Zaragoza-Caballero, 1984
  - Cenophengus wittmeri Zaragoza-Caballero, 1984
- Geslacht Cephalophrixothrix Wittmer, 1976
  - Cephalophrixothrix clypeatus Wittmer, 1976
  - Cephalophrixothrix columbianus Wittmer, 1976
  - Cephalophrixothrix nigerrimus Wittmer, 1976
- Geslacht Decamastinocerus Wittmer, 1988
  - Decamastinocerus parvimandibularis Wittmer, 1988
- Geslacht Distremocephalus Wittmer, 1976
  - Distremocephalus barrerai Zarazoga, 1986
  - Distremocephalus beutelspacheri Zarazoga, 1986
  - Distremocephalus buenoi Zarazoga, 1986
  - Distremocephalus californicus (Van Dyke, 1918)
    - = Mastinocerus californicus Van Dyke, 1918
  - Distremocephalus chiapensis Zarazoga, 1986
  - Distremocephalus leonilae Zarazoga, 1986
  - Distremocephalus mexicanus (Wittmer, 1963)
    - = Mastinocerus mexicanus Wittmer, 1963
  - Distremocephalus opaculus (Horn, 1895)
    - = Mastinocerus opaculus Horn, 1895
  - Distremocephalus rufocaudatus Zarazoga, 1986
  - Distremocephalus texanus (LeConte, 1874)
    - = Mastinocerus texanus LeConte, 1874
  - Distremocephalus wittmeri Zarazoga, 1986
- Geslacht Eurymastinocerus Wittmer, 1976
  - Eurymastinocerus arawakensis Wittmer, 1976
  - Eurymastinocerus columbiana (Wittmer, 1963)
    - = Euryopa columbiana Wittmer, 1963
  - Eurymastinocerus niger Gorham, 1881
    - = Euryopa nigra Gorham, 1881
  - Eurymastinocerus politicollis Wittmer, 1988
  - Eurymastinocerus reductipennis (Wittmer, 1970
    - = Mastinocerus reductipennis Wittmer, 1970
- Geslacht Euryognathus Wittmer, 1976
  - Euryognathus bicoloratus Wittmer, 1976
  - Euryognathus venezolanus (Wittmer, 1956)
    - = Mastinocerus venezolanus Wittmer, 1956
- Geslacht Euryopa Gorham, 1881
  - = Bruchodrilus Pic, 1926
  - Euryopa brasiliensis Wittmer, 1976
  - Euryopa caucaensis Wittmer, 1976
  - Euryopa clarindae Wittmer, 1996
  - Euryopa columbiana Wittmer, 1963
  - Euryopa laurae Wittmer, 1996
  - Euryopa nigra Gorham, 1881
  - Euryopa opacipennis (Pic, 1926)
    - = Bruchodrilus opacipennis Pic, 1926
    - = Euryopa robustior Pic, 1929
  - Euryopa peckorum Wittmer, 1996
  - Euryopa ruficeps Wittmer, 1976
  - Euryopa singularis Gorham, 1881
- Geslacht Howdenia Wittmer, 1976
  - Howdenia coroicoensis Wittmer, 1988
  - Howdenia fischeri (Pic, 1937)
    - = Phrixothrix fischeri Pic, 1937
  - Howdenia frontalis (Wittmer, 1963)
    - = Taximastinocerus frontalis Wittmer, 1963
  - Howdenia golbachi Wittmer, 1988
  - Howdenia minutissima Wittmer, 1988
  - Howdenia neblinensis Wittmer, 1988
  - Howdenia nigerrima Wittmer, 1976
  - Howdenia punctata Wittmer, 1976
  - Howdenia robusta Wittmer, 1988
- Geslacht Mastinocerus Solier in Gay, 1849
  - Ondergeslacht Mastinocerus Solier in Gay, 1849
    - = Mastigocerus Gemminger & Harold, 1869
    - Mastinocerus araucanus Wittmer, 1963
    - Mastinocerus atriceps (Pic, 1915)
      - = Phrixothrix atriceps Pic, 1915
      - = Phrixothrix argentinus Pic, 1915
    - Mastinocerus brevipennis Solier in Gay, 1849
    - Mastinocerus chilensis Wittmer, 1976
    - Mastinocerus germaini Pic, 1930
    - Mastinocerus kuscheli Wittmer, 1976
    - Mastinocerus nigeropacus Wittmer, 1970
    - Mastinocerus nigriceps Wittmer, 1963
    - Mastinocerus nigricollis (Pic, 1915)
      - = Phrixothrix nigricollis Pic, 1915
    - Mastinocerus nigroapicalis Pic, 1915
      - = Phrixothrix nigroapicalis
    - Mastinocerus obscurus Wittmer, 1976
    - Mastinocerus patruelis (Pic, 1915)
      - = Phrixothrix patruelis Pic, 1915
    - Mastinocerus punctatus Wittmer, 1963
    - Mastinocerus ramosus (Gorham, 1881)
      - = Ptorthodius ramosus Gorham, 1881
    - Mastinocerus ruficollis Wittmer, 1976
    - Mastinocerus travassosi Wittmer, 1963
    - Mastinocerus uruguayensis (Berg, 1886
      - = Phengodes uruguayensis Berg, 1886
  - Ondergeslacht Mastinocerus incertae
    - Mastinocerus callanganus Wittmer, 1963
  - Ondergeslacht Paramastinocerus Wittmer, 1976
    - Mastinocerus janeirensis Wittmer, 1976
    - Mastinocerus ondulatus Wittmer, 1963
    - Mastinocerus pauloensis Wittmer, 1963
    - Mastinocerus peruanus (Wittmer, 1976)
      - = Matinocerus peruanus Wittmer, 1976
- Geslacht Mastinomorphus Wittmer, 1976
  - Mastinomorphus atacamensis (Wittmer, 1963)
    - = Mastinocerus atacamensis Wittmer, 1963
  - Mastinomorphus impressiceps Wittmer, 1997
  - Mastinomorphus martinezi (Wittmer, 1963)
    - = Mastinocerus martinezi Wittmer, 1963
  - Mastinomorphus minutus Wittmer, 1976
  - Mastinomorphus misionensis (Wittmer, 1950)
    - = Phrixothrix misionensis Wittmer, 1950
  - Mastinomorphus obscuripennis Wittmer, 1976
  - Mastinomorphus pampaensis (Wittmer, 1950)
    - = Phrixothrix pampaensis Wittmer, 1950
  - Mastinomorphus piceipennis (Pic, 1929)
    - = Phrixothrix piceipennis Pic, 1929
  - Mastinomorphus rufescens (Pic, 1938)
    - = Phrixothrix rufescens Pic, 1938
  - Mastinomorphus ruficeps (Pic, 1926)
    - = Phrixothrix ruficeps Pic, 1926
  - Mastinomorphus vicunaensis Wittmer, 1976
  - Mastinomorphus weiseri (Pic, 1926)
    - = Phrixothrix weiseri Pic, 1926
- Geslacht Mastinowittmerus Zaragoza, 1984
  - Mastinowittmerus mexicanus Zaragoza, 1984
  - Mastinowittmerus vazquezae Zarazoga, 1986
- Geslacht Neophengus Wittmer, 1976
  - Neophengus chilensis Wittmer, 1976
  - Neophengus huantaensis Wittmer, 1976
  - Neophengus nanus (Wittmer, 1948)
    - = Phrixothrix nanus Wittmer, 1948
  - Neophengus penai (Wittmer, 1963)
    - = Cenophengus penai Wittmer, 1963
- Geslacht Nephromma Wittmer, 1976
  - Nephromma alvarengai Wittmer, 1996
  - Nephromma barberi (Wittmer, 1976)
    - = Nephroma barberi Wittmer, 1976
- Geslacht Oxymastinocerus Wittmer, 1963
  - Oxymastinocerus bridarolli Wittmer, 1963
  - Oxymastinocerus fulvus (Philippi, 1864)
    - = Mastinocerus fulvus Philippi, 1864
  - Oxymastinocerus nigripennis Wittmer, 1988
  - Oxymastinocerus pecki Wittmer, 1976
  - Oxymastinocerus peruanus (Wittmer, 1988)
    - = Mastinocerus peruanus Wittmer, 1956
  - Oxymastinocerus rufotestaceus Wittmer, 1976
  - Oxymastinocerus unicolor (Pic, 1926)
    - = Phrixothrix unicolor Pic, 1926
  - Oxymastinocerus venezolanus Wittmer, 1976
- Geslacht Paraphrixothrix Zaragoza-Caballero, 2010^[8]
  - Paraphrixothrix ecuadoranus Zaragoza-Caballero, 2010
- Geslacht Paraptorthodius Schaeffer, 1904
  - Paraptorthodius mirabilis Schaeffer, 1904
  - Paraptorthodius schaefferi (Zaragoza, 1989:88)
    - = Paratorthodius schaefferi Zaragoza, 1989
- Geslacht Phrixothrix Olivier, 1909
  - Phrixothrix acuminatus Pic, 1929
  - Phrixothrix alboterminatus Wittmer, 1963
  - Phrixothrix belemensis Wittmer, 1976
  - Phrixothrix clypeatus Wittmer, 1992
  - Phrixothrix gibbosus Wittmer, 1976
  - Phrixothrix heydeni Olivier, 1910
  - Phrixothrix hieronymi (Haase, 1886)
    - = Phengodes hieronymi Haase, 1886
    - = Phrixothrix obscuripes Pic, 1915
  - Phrixothrix hirtus Olivier, 1909
  - Phrixothrix microphthalmus Wittmer, 1976
  - Phrixothrix obscurus Pic, 1915
  - Phrixothrix peruanus Wittmer, 1963
  - Phrixothrix pickeli Pic, 1933
  - Phrixothrix reducticornis Wittmer, 1963
  - Phrixothrix staphylinoides Wittmer, 1963
  - Phrixothrix tiemanni Wittmer, 1970
  - Phrixothrix vianai Wittmer, 1988
  - Phrixothrix vivianii Wittmer, 1992
- Geslacht Pseudomastinocerus Wittmer, 1963
  - Pseudomastinocerus chiriquiensis Wittmer, 1963
  - Pseudomastinocerus costaricanus Wittmer, 1988
  - Pseudomastinocerus freyi Wittmer, 1963
  - Pseudomastinocerus laticeps (Pic, 1937)
    - = Phrixothrix laticeps Pic, 1937
  - Pseudomastinocerus panamensis Wittmer, 1976
  - Pseudomastinocerus ruficeps (Pic, 1925)
    - = Phengodes ruficeps Pic, 1925
  - Pseudomastinocerus tinalandicus Wittmer, 1976
- Geslacht Ptorthodiellus Wittmer, 1976
  - Ptorthodiellus araguaicus Wittmer, 1976
  - Ptorthodiellus trinidadicus Wittmer, 1976
- Geslacht Ptorthodius Gorham, 1881
  - - = Cephalodrilus Pic
  - Ptorthodius atricornis (Pic, 1929)
    - = Phrixothrix atricornis Pic, 1929
  - Ptorthodius lanei Wittmer, 1963^[9]
  - Ptorthodius mandibularis Gorham, 1881
    - = Cephalodrilus atrosignatus Pic
  - Ptorthodius quiguasensis Wittmer, 1996
- Geslacht Spangleriella Wittmer, 1988
  - Spangleriella vittata Wittmer, 1988
- Geslacht Stenophrixothrix Wittmer, 1963
  - Stenophrixothrix bogotensis (Pic, 1925)
    - = Phengodes bogotensis Pic, 1925
  - Stenophrixothrix chiriquensis Wittmer, 1988
  - Stenophrixothrix curticollis (Pic, 1925)
    - = Phengodes curticollis Pic, 1925
  - Stenophrixothrix darlingtoni Wittmer, 1976
  - Stenophrixothrix ecuadoranus Wittmer, 1988
  - Stenophrixothrix elongatus Wittmer, 1988
  - Stenophrixothrix espiritensis Wittmer, 1996
  - Stenophrixothrix fuscus Gorham, 1881
    - = Euryopa fusca Gorham, 1881
  - Stenophrixothrix howdeni Wittmer, 1988
  - Stenophrixothrix longipes Wittmer, 1976
  - Stenophrixothrix mandibularis Wittmer, 1988
  - Stenophrixothrix martinezi Wittmer, 1988
  - Stenophrixothrix minor Wittmer, 1976
  - Stenophrixothrix montanus Wittmer, 1976
  - Stenophrixothrix nigripennis Wittmer, 1976
  - Stenophrixothrix oculatus Wittmer, 1988
  - Stenophrixothrix pallens (Berg, 1885:232)
    - = Phengodes pallens Berg, 1885
  - Stenophrixothrix panamensis Wittmer, 1988
  - Stenophrixothrix pseudomandibularis Wittmer, 1988
- Geslacht Taximastinocerus Wittmer, 1963
  - Taximastinocerus beniensis Wittmer, 1976
  - Taximastinocerus brasiliensis Wittmer, 1976
  - Taximastinocerus breviplumatus Wittmer, 1988
  - Taximastinocerus brunneus Gorham, 1881
    - = Euryopa brunea Gorham, 1881
  - Taximastinocerus cephalotes (Pic, 1938)
    - = Phrixothrix cephalotes Pic, 1938
  - Taximastinocerus frontalis Wittmer, 1963
  - Taximastinocerus hermanni Wittmer, 1997
  - Taximastinocerus hickeri (Pic, 1929)
    - = Phrixothrix hickeri Pic, 1929
  - Taximastinocerus kissingeri Wittmer, 1963
  - Taximastinocerus levefumidus Wittmer, 1988
  - Taximastinocerus neblinensis Wittmer, 1988
  - Taximastinocerus nigricolor Wittmer, 1988
  - Taximastinocerus pallidus (Pic, 1938)
    - = Phrixothrix pallidus Pic, 1938
  - Taximastinocerus paralleleus Wittmer, 1976
  - Taximastinocerus plaumanni Wittmer, 1963
  - Taximastinocerus pseudobrunneus Wittmer, 1988
  - Taximastinocerus puncticollis Wittmer, 1976
Onderfamilie Penicillophorinae Paulus, 1975
- Geslacht Acladocera Wittmer, 1981
  - Acladocera hispaniolae Wittmer, 1981
- Geslacht Adendrocera Wittmer, 1976
  - Adendrocera flavulum (Wittmer, 1976)
    - = Anendocera flavulum Wittmer, 1976
- Geslacht Penicillophorus Paulus, 1974
  - Penicillophorus ctenotarsus Paulus, 1974
- Geslacht Tarsakanthos Zaragoza-Caballero, 2008^[10]
  - Tarsakanthos minuta Zaragoza-Caballero, 2008^[10]
- Geslacht Walterius Zaragoza-Caballero, 2008^[10]
  - Walterius caballeroae Zaragoza-Caballero, 2008^[10]
  - Walterius emilioi Zaragoza-Caballero, 2008^[10]

Bronnen, noten en/of referenties

↑ V.w.b. onderfamilies Bouchard, P. et al (2011) Family-group names in Coleoptera (Insecta). [[ISSN 1313-2989|ZooKeys]], 88: 1-972. DOI:10.3897/zookeys.88.807 ISBN 978-954-642-583-6 (hardback) ISBN 978-954-642-584-3 (paperback) ZooBank
↑ Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523: 503-504
↑ Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523: 516-518
↑ Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523
↑ Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523: 505-515
↑ Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523: 515
↑ Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523: 516
↑ Zaragoza-Caballero, S. (2010) Paraphrixothrix ecuadoranus, a new genus and species of Phengodidae (Coleoptera: Mastinocerinae) from Ecuador. Pan-Pacific entomologist, 86(4): 105–110. DOI:10.3956/2007-40.1
↑ Wittmer, W. (1963) Zur Kenntnis der Phengodidae (Coleoptera) (25. Beitrag zuer Kenntnis der neotropischen Malacodermata). Mitteilungen der Schweizerischen entomologischen Gesellschaft, 36, 73–99.
↑ ^a ^b ^c ^d ^e Zaragoza-Caballero, S. (2008) Two new genera and three new species of Penicillophorinae (Coleoptera: Phengodidae) of Mexico. Revista mexicana de biodiversidad, 79(2): 363–368. PDF

[1] V.w.b. onderfamilies Bouchard, P. et al (2011) Family-group names in Coleoptera (Insecta). [[ISSN 1313-2989|ZooKeys]], 88: 1-972. DOI:10.3897/zookeys.88.807 ISBN 978-954-642-583-6 (hardback) ISBN 978-954-642-584-3 (paperback) ZooBank

[2] Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523: 503-504

[3] Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523: 516-518

[4] Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523

[5] Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523: 505-515

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[7] Roza, A.S., Quintino, H.Y.S., Mermudes, J.R.M & Silveira, Luiz F.L. Da (2017) Akamboja gen. nov., a new genus of railroad-worm beetle endemic to the Atlantic Rainforest, with five new species (Coleoptera: Phengodidae, Mastinocerinae), Zootaxa 4306 (4), pp. 501-523: 516

[8] Zaragoza-Caballero, S. (2010) Paraphrixothrix ecuadoranus, a new genus and species of Phengodidae (Coleoptera: Mastinocerinae) from Ecuador. Pan-Pacific entomologist, 86(4): 105–110. DOI:10.3956/2007-40.1

[9] Wittmer, W. (1963) Zur Kenntnis der Phengodidae (Coleoptera) (25. Beitrag zuer Kenntnis der neotropischen Malacodermata). Mitteilungen der Schweizerischen entomologischen Gesellschaft, 36, 73–99.

[Zaragoza2008-10] Zaragoza-Caballero, S. (2008) Two new genera and three new species of Penicillophorinae (Coleoptera: Phengodidae) of Mexico. Revista mexicana de biodiversidad, 79(2): 363–368. PDF

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